PhD, University of California, Berkeley

PhD, National Chung Hsing University Institute of BioTechnology

PhD, University of Edinburgh, Scotland

PhD, Institute of Biopharmaceutical Sciences, National Yang-Ming University

Ph.D. University of Illinois at Urbana-Champaign, USA

_{Molecular characteristics and functions of Circular RNAs.}

Ph.D. Institute of Bioinformatics and structural Biology, National Tsing Hua University

Structural biology

PhD in Plant Cell Biology, Wageningen University & Research (WUR)

_{Mechanism of Plant Stem Cell Regeneration}

Ph.D, Institute of Chemistry and Biochemistry, Freie Universität Berlin (Germany)

Structure biology in cellular signaling and membrane trafficking